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<article article-type="research-article" dtd-version="1.3" xmlns:mml="http://www.w3.org/1998/Math/MathML" xmlns:xlink="http://www.w3.org/1999/xlink" xmlns:xsi="http://www.w3.org/2001/XMLSchema-instance" xml:lang="ru"><front><journal-meta><journal-id journal-id-type="publisher-id">vavilov</journal-id><journal-title-group><journal-title xml:lang="ru">Вавиловский журнал генетики и селекции</journal-title><trans-title-group xml:lang="en"><trans-title>Vavilov Journal of Genetics and Breeding</trans-title></trans-title-group></journal-title-group><issn pub-type="epub">2500-3259</issn><publisher><publisher-name>Institute of Cytology and Genetics of Siberian Branch of the RAS</publisher-name></publisher></journal-meta><article-meta><article-id pub-id-type="doi">10.18699/VJ19.471</article-id><article-id custom-type="elpub" pub-id-type="custom">vavilov-1923</article-id><article-categories><subj-group subj-group-type="heading"><subject>Research Article</subject></subj-group><subj-group subj-group-type="section-heading" xml:lang="ru"><subject>ОРГАНИЗАЦИЯ ХРОМОСОМ</subject></subj-group><subj-group subj-group-type="section-heading" xml:lang="en"><subject>CHROMOSOME ORGANIZATION</subject></subj-group></article-categories><title-group><article-title>Особенности молекулярного состава гетерохроматина, ассоциированного с проядрышками в эмбрионах мыши</article-title><trans-title-group xml:lang="en"><trans-title>Peculiarities of the molecular composition  of heterochromatin associated with pronucleoli  in mouse embryos</trans-title></trans-title-group></title-group><contrib-group><contrib contrib-type="author" corresp="yes"><name-alternatives><name name-style="eastern" xml:lang="ru"><surname>Боголюбова</surname><given-names>И. О.</given-names></name><name name-style="western" xml:lang="en"><surname>Bogolyubova</surname><given-names>I. O.</given-names></name></name-alternatives><bio xml:lang="ru"><p>Санкт-Петербург.</p></bio><bio xml:lang="en"><p>St. Petersburg.</p></bio><email xlink:type="simple">ibogol@mail.ru</email><xref ref-type="aff" rid="aff-1"/></contrib><contrib contrib-type="author" corresp="yes"><name-alternatives><name name-style="eastern" xml:lang="ru"><surname>Сайлау</surname><given-names>Ж. К.</given-names></name><name name-style="western" xml:lang="en"><surname>Sailau</surname><given-names>Z. K.</given-names></name></name-alternatives><bio xml:lang="ru"><p>Санкт-Петербург.</p></bio><bio xml:lang="en"><p>St. Petersburg.</p></bio><xref ref-type="aff" rid="aff-2"/></contrib><contrib contrib-type="author" corresp="yes"><name-alternatives><name name-style="eastern" xml:lang="ru"><surname>Боголюбов</surname><given-names>Д. С.</given-names></name><name name-style="western" xml:lang="en"><surname>Bogolyubov</surname><given-names>D. S.</given-names></name></name-alternatives><bio xml:lang="ru"><p>Санкт-Петербург.</p></bio><bio xml:lang="en"><p>St. Petersburg.</p></bio><xref ref-type="aff" rid="aff-1"/></contrib></contrib-group><aff-alternatives id="aff-1"><aff xml:lang="ru">Институт цитологии Российской академии наук.<country>Россия</country></aff><aff xml:lang="en">Institute of Cytology, RAS.<country>Russian Federation</country></aff></aff-alternatives><aff-alternatives id="aff-2"><aff xml:lang="ru">Санкт-Петербургский государственный университет.<country>Россия</country></aff><aff xml:lang="en">St. Petersburg State University.<country>Russian Federation</country></aff></aff-alternatives><pub-date pub-type="collection"><year>2019</year></pub-date><pub-date pub-type="epub"><day>28</day><month>03</month><year>2019</year></pub-date><volume>23</volume><issue>2</issue><fpage>129</fpage><lpage>134</lpage><permissions><copyright-statement>Copyright &amp;#x00A9; Боголюбова И.О., Сайлау Ж.К., Боголюбов Д.С., 2019</copyright-statement><copyright-year>2019</copyright-year><copyright-holder xml:lang="ru">Боголюбова И.О., Сайлау Ж.К., Боголюбов Д.С.</copyright-holder><copyright-holder xml:lang="en">Bogolyubova I.O., Sailau Z.K., Bogolyubov D.S.</copyright-holder><license license-type="creative-commons-attribution" xlink:href="https://creativecommons.org/licenses/by/4.0/" xlink:type="simple"><license-p>This work is licensed under a Creative Commons Attribution 4.0 License.</license-p></license></permissions><self-uri xlink:href="https://vavilov.elpub.ru/jour/article/view/1923">https://vavilov.elpub.ru/jour/article/view/1923</self-uri><abstract><p>Ядро доимплантационных эмбрионов млекопитающих характеризуется своеобразной структурной организацией. На начальных стадиях дробления вместо функционально активных ядрышек в ядре эмбриона присутствуют так называемые проядрышки – фибриллярные электронно-плотные структуры, неактивные в отношении синтеза РНК. Подавляющее большинство проядрышек окружено кольцеообразной зоной транскрипционно неактивного гетерохроматина, который, однако, содержит не только эпигенетическую метку репрессированного хроматина – триметилированный гистон Н3K9me3, но и метку активного хроматина – ацетилированный гистон H4K5ac. По результатам непрямого иммуномечения, молекулярный состав кольцевого гетерохроматина в эмбрионах мыши изменяется в ходе реализации процессов активации эмбрионального генома, а также при искусственном подавлении транскрипционной активности. На стадии зиготы в составе кольцевого гетерохроматина выявляются некоторые факторы метаболизма мРНК, например SR-белок фактор сплайсинга SC35 (SRSF2) и базальный фактор транскрипции TFIID. На более поздних стадиях развития в этой области начинают выявляться другие ядерные белки, например Y14 – коровый компонент комплекса связи экзонов (EJC), а также белки, вовлеченные в ремоделинг хроматина – ATRX и Daxx. Типичным компонентом кольцевого гетерохроматина является актин, иммуноцитохимическое мечение которого наиболее выражено на двухклеточной стадии дробления, после активации эмбрионального генома. Характерно, что молекулярный состав гетерохроматина, ассоциированного с разными проядрышками в одном ядре, может различаться, что, возможно, отражает функциональную гетерогенность морфологически сходных проядрышек по их компетентности к процессу нуклеологенеза. В настоящем обзоре кратко обсуждаются некоторые особенности молекулярного состава гетерохроматина, ассоциированного с NPBs, и его возможные функции.</p></abstract><trans-abstract xml:lang="en"><p>The nucleus of pre-implantation mammalian embryos is characterized by peculiar structural organization. At the initial stages of cleavage, the nucleus of the embryo contains the so-called nucleolus precursor bodies (NPBs) or pronucleoli rather than functionally active nucleoli. The NPBs are fibrillar electron-dense structures inactive in RNA synthesis. The vast majority of NPBs are surrounded by a ring-shaped zone of transcriptionally inactive heterochromatin. Intriguingly, these zones contain not only tri-methylated histone Н3K9me3 as an epigenetic mark of repressed chromatin but also acetylated histone H4K5ac, a well-known marker of active chromatin. Immunocytochemical data suggest that the molecular composition of this ‘ring heterochromatin’ in mouse embryos changes during the realization of embryonic genome activation events, as well as during artificial suppression of transcription. In zygotes, some factors of mRNA biogenesis including splicing factor SC35 (SRSF2) and basal transcription factor TFIID are detectable in the ring chromatin. At later stages of development, other nuclear proteins such as Y14, a core component of the exon-exon junction complex (EJC), as well as the proteins involved in chromatin remodeling (ATRX, Daxx) are also detectable in this area. A typical component of the ‘ring heterochromatin’ is actin. Anti-actin immunocytochemical labeling is most expressed at the two-cell cleavage stage after activation of the embryonic genome. Indicatively, the molecular composition of the ‘ring heterochromatin’ associated with different NPBs may differ significantly even in the same nucleus. This seems to reflect the functional heterogeneity of morphologically similar NPBs according to their competence to the process of nucleologenesis. Here, we discuss briefly some peculiarities of the molecular composition and possible functions of the NPB-associated heterochromatin in mouse early embryos.</p></trans-abstract><kwd-group xml:lang="ru"><kwd>гетерохроматин</kwd><kwd>доимплантационные эмбрионы мыши</kwd><kwd>иммуноцитохимия</kwd></kwd-group><kwd-group xml:lang="en"><kwd>heterochromatin</kwd><kwd>pre-implantation mouse embryos</kwd><kwd>immunocytochemistry</kwd></kwd-group></article-meta></front><back><ref-list><title>References</title><ref id="cit1"><label>1</label><citation-alternatives><mixed-citation xml:lang="ru">Banani S.F., Lee H.O., Hyman A.A., Rosen M.K. Biomolecular condensates: organizers of cel-lular biochemistry. Nat. Rev. Mol. Cell Biol. 2017;18(5):285-298. DOI 10.1038/nrm.2017.7. Bogolyubova I.O. Transcriptional activity of nuclei in 2-cell blocked mouse embryos. Tissue Cell. 2011;43(4):262-265. 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